The marsh harriers are

birds of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators ...

of the harrier subfamily. They are medium-sized

raptor Raptor or RAPTOR may refer to: Animals The word "raptor" refers to several groups of bird-like dinosaurs which primarily capture and subdue/kill prey with their talons. * Raptor (bird) or bird of prey, a bird that primarily hunts and feeds on ...

s and the largest and broadest-winged harriers. Most of them are associated with marshland and dense reedbeds. They are found almost worldwide, excluding only the

Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...

. Until recently two species were generally recognized: the marsh harrier (''Circus aeruginosus'') and the

African marsh harrier The African marsh harrier (''Circus ranivorus'') is a bird of prey belonging to the harrier genus ''Circus''. It is largely resident in wetland habitats in southern, central and eastern Africa from South Africa north to South Sudan. Description ...

(''C. ranivorus''). The marsh harrier is now usually split into several species, sometimes as many as six. These are the

western marsh harrier The western marsh harrier (''Circus aeruginosus'') is a large harrier, a bird of prey from temperate and subtropical western Eurasia and adjacent Africa. It is also known as the Eurasian marsh harrier. Formerly, a number of relatives were includ ...

(''C. aeruginosus''),

eastern marsh harrier The eastern marsh harrier (''Circus spilonotus'') is a bird of prey belonging to the marsh harrier group of harriers. It was previously considered to be conspecific with the western marsh harrier (''Circus aeruginosus'') but is now usually classi ...

(''C. spilonotus''),

Papuan harrier The Papuan harrier (''Circus spilothorax'') is a bird of prey native to New Guinea which belongs to the harrier genus ''Circus''. It was classified as a subspecies of the eastern marsh harrier (''Circus spilonotus'') of eastern Asia. It is thi ...

(''C. spilonotus spilothorax'' or ''C. spilothorax''),

swamp harrier The swamp harrier (''Circus approximans''), also known as the Australasian marsh harrier, Australasian harrier or swamp-hawk, is a large, slim bird of prey widely distributed across Australasia. In New Zealand it is also known as the harrier haw ...

(''C. approximans''),

Réunion harrier The Réunion harrier or Réunion marsh harrier (''Circus maillardi'') is a bird of prey belonging to the marsh harrier group of harriers. It is now found only on the Indian Ocean island of Réunion, although fossil material from Mauritius has be ...

(''C. maillardi maillardi'' or ''C. maillardi'') and

Madagascar marsh harrier The Malagasy harrier (''Circus macrosceles'') is a bird of prey belonging to the marsh harrier group of harriers. It inhabits Madagascar and the Comoro Islands in the Indian Ocean. It was formerly regarded as a subspecies of the Réunion harrie ...

(''C. maillardi macrosceles'' or ''C. macrosceles''). At the beginning of the 20th century, the marsh harrier was hunted to extinction in the

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...

. After being reintroduced from other regions, its population steadily increased until

DDT Dichlorodiphenyltrichloroethane, commonly known as DDT, is a colorless, tasteless, and almost odorless crystalline chemical compound, an organochloride. Originally developed as an insecticide, it became infamous for its environmental impacts. ...

threatened it, along with other raptors, in the 1950s and 1960s. Since, the population has slowly and steadily increased.

Species

The

(''Circus aeruginosus''), often simply called the marsh harrier, breeds widely across

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

and

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

. It is migratory except in the mildest regions, and winters mainly in

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

. It hunts small

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...

s and

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

s, surprising them as it drifts low over fields and reedbeds. Its long legs allow it to pluck frogs and fish from the water mid-swoop. The western marsh harrier is a typical harrier, with long wings held in a shallow V in its low flight. It also resembles other harriers in having distinct male and female plumages, but its plumages are quite different from those of its relatives. The male has wings with grey and brown sections and black wingtips. Its head, tail and underparts are greyish, except for the chestnut belly. The female is mainly brown with a cream crown and cream leading edge to her wings. It is 19-22 inches long and weighs 1-2 lbs. The

(''C. spilonotus'') breeds in the grasslands and wetlands of southern

Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...

, northern

Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...

, north-east

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

Manchuria Manchuria is an exonym (derived from the endo demonym " Manchu") for a historical and geographic region in Northeast Asia encompassing the entirety of present-day Northeast China (Inner Manchuria) and parts of the Russian Far East (Outer Manc ...

and

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

, and migrates for the northern winter to

South-east Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...

, the

Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...

and northern

Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...

. Like all marsh harriers, it favours open, wet environments, and is frequently seen drifting low over rice fields, interspersing long, watchful circling glides with two or three slow, powerful wingbeats. Two subspecies are recognised: ''C. s. spilonotus'' of east Asia, and ''C. s. spilothorax'', of

New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...

and the islands nearby, which is sometimes thought to be a separate species and is often called the

. The

(''C. approximans'') at about 50 to 58 cm is slightly larger than ''C. spilonotus'', often a little darker, and has less heavily barred wings and tail. It is commonly found in suitable habitat anywhere in

Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecologica ...

, particularly in the higher rainfall areas to the east, south-east, and south-west, of

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

and throughout

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

, but also in the tropical north of Australia and the island groups of

New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...

Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of no ...

Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...

Tonga Tonga (, ; ), officially the Kingdom of Tonga ( to, Puleʻanga Fakatuʻi ʻo Tonga), is a Polynesian country and archipelago. The country has 171 islands – of which 45 are inhabited. Its total surface area is about , scattered over in ...

and the

Society Islands The Society Islands (french: Îles de la Société, officially ''Archipel de la Société;'' ty, Tōtaiete mā) are an archipelago located in the South Pacific Ocean. Politically, they are part of French Polynesia, an overseas country of the F ...

. It is also known as the Australasian harrier or Pacific marsh harrier. The

or Madagascar harrier (''C. maillardi'') is found on the

Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by th ...

islands of

Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...

Réunion Réunion (; french: La Réunion, ; previously ''Île Bourbon''; rcf, label= Reunionese Creole, La Rényon) is an island in the Indian Ocean that is an overseas department and region of France. It is located approximately east of the island ...

and the

Comoros The Comoros,, ' officially the Union of the Comoros,; ar, الاتحاد القمري ' is an independent country made up of three islands in southeastern Africa, located at the northern end of the Mozambique Channel in the Indian Ocean. It ...

. It is sometimes split into two species: Madagascar harrier (''Circus macrosceles'') and Réunion harrier (''C. maillardi). The

(''C. ranivorus'') is a distinct non-migratory species that has not usually been included as a subspecies of ''C. aeruginosus''. It inhabits southern and eastern

Gallery

File:Circus aeruginosus with prey.JPG, Western marsh harrier with prey, India File:Circus aeruginosus MHNT.ZOO.2010.11.82.5.jpg, ''Circus aeruginosus'' - MHNT File:Marsh Harrier Female.jpg, Marsh Harrier female File:MarshHarrier.jpg, alt=, Marsh Harrier in Northern Israel

References

{{Reflist * Roger Clarke (1995) ''The Marsh Harrier'', Hamlyn, London. * James Ferguson-Lees (2001) ''Raptors of the World'', Christopher Helm.

External links

Ageing and sexing (PDF) by Javier Blasco-Zumeta
Harriers (birds) Bird common names